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Sarah, an orphan, gets caught up in a deadly conspiracy after learning that she is a clone. In the opener, Sarah witnesses the suicide of a woman who looks just like her and assumes the victim's identity in hopes of escaping her troubled life.
Sarah continues to impersonate Beth and gets her a second chance on the force.
Sarah's fake detective ruse stretches thin when the police find the body that she buried.
Sarah gets closer to finding the killer, but her efforts threaten to end her chances of a reunion with her daughter.
Evidence suggests that the orphans are part of an ongoing experiment. Sarah suspects Paul, but her probing inadvertently triggers Paul's own investigation.
Alison's paranoia puts all of the orphans at risk. Meanwhile, Sarah must decide who to trust with her secret.
Sarah confronts the conspiracy head-on, while the trust between herself and Paul weakens. Meanwhile, Cosima grows closer to Delphine and the mysterious Dr. Leekie.
Sarah's actions pit the orphans against each other; Paul tries to cover up Sarah's existence; Alison takes vengeance on her suspected monitor.
Dr. Leekie makes Sarah reconsider taking revenge after Kira is threatened. Meanwhile, Cosima faces the truth about Delphine; and Sarah decides the killer's fate.
The Orphans must decide whether to make peace with their creators; Sarah is forced into a showdown with the enemy.
Series 2 begins with Sarah searching for her missing daughter and suspecting that ruthless pro-clone Rachel is behind the disappearance. Meanwhile, Alison attends the funeral of her fallen friend, Aynsley, which causes Alison to sink into despair.
Sarah gets Art to help her find Kira and is shocked when she learns where the trail leads. Meanwhile, Alison struggles with Aynsley's death; and Cosima grapples with the worsening symptoms of her mysterious sickness.
Sarah turns to an old flame from her past as she hits the road with Felix, but their brief respite is brutally shattered. Sarah then realises that no matter how far she runs it will never be far enough.
Sarah comes home to search for answers and with Cosima's help she digs into the origins of the clone experiment. The hunt takes her into perilous territory that could cost her more than she realizes.
Rachel goes on the warpath and lashes out at Sarah's loved ones. While Sarah tries to keep them safe, an unexpected visitor threatens to derail her plans.
Sarah joins forces with an unlikely ally, when she follows a trail of clues that she hopes will lead her to the origins of the clone experiment. Meanwhile, Cosima and Delphine make a discovery that could prove to be the key to treating Cosima's illness.
A rehab confidante betrays Alison and threatens to expose the truth about Aynsley's death, which causes Alison to go into a free fall on the eve of Family Day. The clones gather to contain the fallout.
A new player in the clone conspiracy emerges and turns up at Felix's door, which sends him into crisis mode. Meanwhile, Sarah struggles with whether to surrender a crucial piece of leverage and strike a deal with Dyad.
Cosima takes a turn for the worse, which forces Sarah to take desperate measures that result in tragic consequences for those she loves. Meanwhile, the Proletheans set their ultimate plan in motion; and Alison and Donnie tangle with the law.
The war with Dyad is all but lost when Rachel's latest ploy forces a broken Sarah to concede. The Clone Club springs into action, uniting in a reckless gambit to save Sarah. As they fight to gain their freedom, Sarah encounters a new ally and uncovers a terrifying new dimension to the conspiracy.
Season 3 begins with Sarah pursuing the missing Helena who is being held in a faraway compound. Sarah uses her wiles and acting chops to deter a Topside investigator who threatens the Leda sisters' lives.
Castor pursues the Leda clones for original tissue samples. Meanwhile, Helena, who is pregnant and imprisoned, is examined by a mysterious villain.
Leda and Castor are both on the hunt for original clone DNA. On the lam, Mark and Gracie pursue new leads to find the samples that might set them free, while Art and Sarah reunite to track them down. Meanwhile, Cosima and Scott try to learn more about Castor's biology and Alison's new money-making venture is going gangbusters. A shocking revelation about the Castor-Leda connection leaves the sisters reeling, just before Sarah witnesses a deadly Prolethean-Castor face-off.
Mark is taken down in a Prolethean shoot-out and Sarah must step in to help. But as Sarah gathers intel that puts her on the trail of the original Castor DNA, a familiar foe is not far behind. While Helena continues to toil in a faraway military compound, it is ultimately Sarah's fate that will hang in the balance as she tries to save her sister. Meanwhile, Gracie is cast out from the Prolethean fold, and Alison and Donnie come face-to-face with unhappy drug lord, Jason Kellerman.
Helena and her seestra are reunited at the Castor camp when Sarah is captured. As Sarah suffers testing, Paul appears to chide her; but his allegiances are beginning to strain. While the Leda family worries about Sarah's absence, Gracie ‘breaks Amish' under the care of Mrs. S and an unwilling Felix. Cosima falls hard for an internet date, Shay.
Paul is pushed into action when he discovers the ugly truth behind the Castor Boys' logbooks of names. Meanwhile, Felix becomes desperate for information on Sarah's whereabouts and presses the still-ailing Rachel for any shred of a lead.
Alison must secure a store front in order to take her and Donnie's business to the "next level." But things don't go according to plan when her mother, Connie, proves to be a major hindrance. Cosima is facing familiar health issues and calls upon beleaguered Alison for a favor. But with Alison working to save an imperiled Donnie from the merciless hands of a drug lord, it's Cosima who ends up doing a favor. An unassuming Scott gets closer to the greatest discovery of all, when he sits down with beleaguered clone Rachel.
When Rachel makes a promise to Sarah, it comes with a hefty price. Meanwhile, Helena and Gracie work with the Hendrixes, and Donnie takes Kellerman to task. Over at Castor, Coady and Benchman scheme to track down the Castor Original, and Rudy descends on DYAD for a much needed asset.
Clone Club heads to London to track down the Castor Original, but their enemies follow closely.
Kendall may be the key to stop Castor; the sisters appear to be thwarting their most dangerous foe.
After two months of respite, Sarah's hard-won refuge in Iceland is shattered by a Neolution attack. Once again forced to flee, she realizes no matter how far her family runs it will never be far enough.
After evading a Neolution attack in Iceland, Sarah returns home to track down an elusive new ally. She follows breadcrumbs and uncovers the next facet of Neolution's agenda – implanting dangerous technological devices in humans. However, those she left behind are wary about her return, not wanting to relinquish their semblance of normalcy. As Cosima struggles with Delphine's disappearance, Alison and Donnie host a pregnant and ever-hungry Helena. Meanwhile Felix, feeling like an outsider, embarks on his own journey of self-discovery.
As Sarah reels from the shocking discovery of Neolution's biotech, she's desperate to find answers. Always impulsive, she follows a lead into a harrowing encounter that requires the help of a reluctant new player. Eager to assist, Mrs. S strikes an alliance with an old nemesis. Rachel receives a visit from her keeper and takes a fraught step forward towards recovery. Meanwhile, Helena is mistaken for "Mrs. Hendrix" when she answers a visit from suspicious visitors.
Sarah forms an alliance with a former enemy in hopes of getting to the head of Neolution. Elsewhere: Alison orders Felix and Donnie to invade a Neolution fertility clinic and a Susan Duncan leaves Rachel to make a devastating decision.
While Sarah takes the day off to reconnect with Kira, Cosima steps up and infiltrates an exclusive Neolutionist fertility research center. A shocking discovery of their clandestine experiments leads her into a dangerous encounter.
When Cosima admits doubts about finding a cure in time to save her life, Sarah makes a lone decision that exposes Kendall to those she's spent years hiding from. Meanwhile, Art and Felix team up to protect naïve clone, Krystal Goderitch, from herself.
Sarah regresses to old habits while struggling to cope with a loss. Following in Beth's footsteps, Sarah questions her strength to carry on.
As Alison is terrified by Neolution threats made against Donnie, Sarah and Mrs. S question her loyalty to her sestras. Siobhan sneaks out to seek vengeance. Cosima and Susan Duncan work together in a final attempt to discover a cure.
With Susan Duncan deposed, Sarah finds herself in the same position as Rachel–under Evie Cho's heel. The archenemies form an uneasy alliance in an attempt to bring Evie down and restore Susan to power.
When communication with Cosima is cut off and Neolution appears to be within striking distance, Sarah's alliance with Rachel goes out the window. Sarah lures a high-level Neolutionist into a trap, and sets out to bring down Neolution once and for all.
Sarah is deeply wounded and struggling to stay alive on the Island. Driven deep into its forest by Neolutionists and worse, she strives to make it far enough to rescue Cosima. Meanwhile, a recovering Cosima reunites with Delphine, but there's way more going on this Island than Cosima imagined and Delphine can't give her all the answers she needs. What is clear is that Rachel is now in firmly in charge at the right-hand of the 170-year-old founder of Neolution, PT Westmorland. Back home, Neolution puts the screws to Alison and Donnie in their hunt for Helena. The sestras couldn't be further apart when they need most to be together.
Captured by Neolution, Sarah receives a damning ultimatum from Rachel: come into the fold, or lose your family. Her decision causes a clash with Kira who begins exploring her own maturity. Meanwhile, Ferdinand struggles to get a handle on Rachel's new velvet glove mandate in pursuit of the clones. With Neolutionists and Ferdinand watching their every move, Felix brings an old ally into the mix to help his family get Kira to safety. On the Island, Cosima struggles to comprehend the reality of the Neolution myth made real.
Stuck on the sidelines with Neolutionists crawling all over her home, Alison seeks to return to her community center, but Bailey Downs has moved on from her. As she flounders to take back some semblance of control, the ghost of friendships' past rears its handsome head. Art and his new partner show up at the Hendrix residence to continue their search for Helena, forcing Art to make a difficult moral judgement.
Mrs. S. and Sarah set out to investigate a lead on a Neolution defector; an unexpected reunion takes place; Cosima has a terrible discovery.
Sarah and Kira spend time at home; secrets are unlocked about Kira's gifts; Cosima joins Delphine at the mansion.
Cosima is forced into an alliance that accelerates P.T. Westmorland's agenda; Krystal stumbles onto a vital clue.
Westmoreland demands a more aggressive action on Kira; Sarah fights to save her daughter; Rachel discovers a dark secret; Cosima has a clue about Westmoreland's identity.
Mrs. S orders the sisters to celebrate Felix's art opening; Sarah remains on edge with a nagging suspicion that S is hiding something.
A devastating loss is suffered; Sarah learns that Helena has been kidnapped; Clone Club bands together to find Helena before her babies are born.
Helena takes shelter behind enemy lines, and Sarah and Art struggle to protect her and end the siege.