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In the premiere episode, "Pilot," an image of Anna, the leader of the V's, is projected worldwide as she speaks about everyone joining together and no longer being divided by country or separated by fear. She wants us all to unite, and is counting on a very important component of human nature -- devotion. At first considered a threat, the V's quickly become a fascination and a link to things that lay just beyond our reach. For Tyler, a teenaged boy, the V's are his ticket to being a part of something big and something that offers hope. To Chad, an egotistical news anchor who wants to further his career, his exclusive interview with Anna begins his desire to dominate the airwaves. Father Jack, a priest with questioning faith, is hesitant to believe in the Visitors' righteousness and goes outside of the church in search of the truth. There are other dissidents who believe the Visitors aren't who they say they are, including Ryan, who is faced with a life-altering decision when the V's show up. And to FBI Counter Terrorist Agent Erica Evans, who discovers, while investigating a terrorist cell, what lurks beneath the alarmingly human exterior of the Visitors, resisting this new world has never been more important -- and never has there been more at stake. This truly is the dawning of a new day.

Erica and Father Jack, having quickly determined they're both against the arrival of the Visitors, suddenly find themselves being tracked by a "Seeker" from the V's. Meanwhile Chad, thinking he blew it for 80 million viewers with his exclusive first interview with Anna, looks to pick himself up and so becomes more investigative in his next newscast; and Dale Maddox's wife and law enforcement begin questioning Erica about his whereabouts.

Chad reports from the Peace Ambassador Center as 100 diplomatic visas are being issued to the first wave of American Visitors, with Anna getting the 1st, but not everyone agrees with the decision. Meanwhile Erica has started tracking a death threat while paired with a V officer, as she actually has to protect the V's, and Ryan starts reaching out to his old friends to build up opposition forces and help fight-off the V's.

Erica works with newly-formed allies to uncover a biological threat they suspect the Visitors have been plotting. Aboard the Mothership, Anna meets with a special guest while managing the investigation into the murder of a V. And Chad does a segment on the V Healing Centers, demonstrating their amazing medical abilities, but then finds himself conflicted by some of his findings.

As the story continues, Erica finds her life in danger when she's attacked at home; a dangerous new member is recruited for the Resistance, as the Visitors shore up their defenses; Anna wonders why Chad hasn't followed up on his diagnosed aneurysm; and Ryan's concern grows over Val's "abnormal" pregnancy.

Anna introduces her pilot program to invite select groups of humans to stay on each of the 29 space ships, Ryan must deal with the fact that Valerie's pregnancy is wildly accelerated, Tyler learns a secret about his mom, and one of the team makes a sacrifice to save Ryan.

Erica, Ryan and Jack go on a harrowing mission to find the legendary John May in order to rescue Georgie, Anna brings Chad aboard the Mothership for a story on her Live Aboard Program, and Tyler confronts his mom about his mysterious past.

Chad and Anna are in Geneva, Switzerland for the U.N. Energy Summit, where Anna is going to present a gift of technology to the world. Meanwhile, Erica learns that the V Task Force is investigating The Fifth Column, and Valerie goes on the run, knowing something is different about her baby and not sure she can trust Ryan.

Erica, Father Jack and Hobbes realize that the Vs know names and addresses of The Fifth Column members and must take drastic measures to protect them. Meanwhile, Ryan finally reveals to Val that he's a Visitor, and Chad, with the camera rolling for "Prime Focus," begins the process aboard the Mothership's Medical Bay to have his aneurysm removed.

Erica, Ryan, Father Jack and Hobbes learn Anna's sent a V shuttle filled with a deadly V tracker team to find them, and must figure out how to stop the shuttle from landing. Meanwhile, Anna gives Tyler his invitation to the Live Aboard Program, and Chad confronts Father Jack on what he knows about The Fifth Column.

After a violent attack on someone very close to Anna, she delivers a message that the Vs don't feel safe and will be leaving; and Anna also gives a list of possible Fifth Column members to Chad, hoping he'll investigate to help in finding them. Meanwhile, Erica and the team try to track down a scientist who may have created a weapon that the Visitors are afraid of.

The episode opens with the V soldier still on the hunt for Val, while for her part, Val's water breaks and her baby with Ryan will soon be born. Meanwhile Erica, tired of being on the defensive, decides to go on the offensive and gets herself and Tyler invited up to the Mothership for a get-to-know-everyone-better dinner. But she has more than food on her mind; she's really there to find Anna's soldiers and destroy them. And Chad, after Father Jack asks him to help get a message on the ship to Joshua, discovers the awful truth that Anna's been using him.

The Season Two opener, "Red Rain," continues the action from last season's finale when Anna unleashed Red Sky on the planet. With the world on the brink of war and chaos all around, is there anyone who can stand up to Anna? Meanwhile, Ryan has been kept in isolation on the mothership, away from his hybrid baby so that Anna can experiment on her; Chad, seeking redemption for being used by Anna, now wants to fight with the Fifth Column; and Erica goes on a journey to find a scientist who can give her the answers to what Red Sky is, and where she learns the horrifying secret of what the V's are doing on our planet. Jane Badler and Bret Harrison join the series in recurring roles.

Anna and Diana square off in the dark bowels of the Mothership - Diana has been down there for 15 years and not had a single glimpse of Anna. Meanwhile, Erica has Tyler's blood analyzed, and learns the V's did something sinister to her when she was pregnant with him.

Erica learns her partner, Malik, is a Visitor and must fight for her life - literally - to stop Anna from finding out she's Fifth Column. Meanwhile, Anna learns that Malik is missing and orders Ryan to find her, while Lisa realizes her body is beginning to change, causing her to become more and more like her mother.

After the radical faction of the Fifth Column murders three Peace Ambassadors, Erica goes on a mission to hunt down their leader, Eli Cohn, a former Mossad agent. Meanwhile, as Jack and other priests around the globe begin to speak out against the Visitors, Anna heads to the Vatican in an effort to convince the leaders of organized religion that the Visitors truly are "of peace," on "V," TUESDAY, FEBRUARY 1 on ABC. Also in the episode, Jay Karnes guest stars as Chris Bolling, an FBI agent now assigned to work with Erica; while Oded Fehr will play Eli Cohn, the leader of a much more radical group opposed to the Visitors.

Anna uses her newest gift to mankind, "Concordia," as a cover to hide an invading fleet of breeding vessels that will be used to ensure the perpetuation of Visitors for future generations. Meanwhile, Eli Cohn tries to convince Erica and the Fifth Column that the "Concordia" gala is the perfect time to assassinate Anna.

Anna, enraged at the increasing audacity of the Fifth Column attacks, enlists Ryan in an attempt to wipe out the Fifth Column in one fell swoop. Meanwhile, Father Travis, following Vatican orders, asks Jack to leave the church, and Lisa meets Diana, her grandmother -- who Anna said was dead -- for the first time.

Erica, seeking revenge against Anna, heads to Hong Kong to carry out a mission to strike a lethal blow against Anna. Meanwhile, Anna learns that Tyler's phosphorous level is too low - even after the Red Rain -- and he might become useless to her; and Lisa and Diana bond, beginning to form an alliance against Anna.

Erica, now officially in charge of the global Fifth Column, leads them in a high risk attack against Anna. Anna meanwhile, growing closer in her plan to destroy humanity, finds a major obstacle in a certain Live Aboard candidate; and Ryan sneaks back onto the Mothership to try and rescue his daughter.

Anna is about to pull the switch on the planet's first Blue Energy reactor, but Sid realizes that the size of the reactor is much too large for just the various Concordia sites - the Fifth Column needs to figure out what Anna is really up to. And Diana tells Lisa she must act against Anna now, fulfill her destiny and save humans from annihilation.

Anna is focused on having Lisa breed with Tyler to ensure the evolution of the Visitors, while Erica and Diana enlist Lisa to stage a coup against Anna and remove her from power before humanity is destroyed. Erica and the Fifth Column join Lisa in the attempted coup, which has devastating results, on the shocking Season Two finale of "V," TUESDAY, MARCH 15 on ABC. Also in the episode, Marc Singer, who played freedom fighter Mike Donovan on the original "V" series, will guest star as Lars Tremont, a member of a top secret organization of high-ranking military and government leaders from around the globe who have long suspected that the Visitors are very much not here in peace.